In tomato, bacterial speck disease (caused by Pseudomonas syringae pv. tomato) is both an economically important disease and an excellent model system to study plant immunity and molecular biology in a crop species.  Speck disease causes necrotic lesions on fruit, leaves, flowers, and stems which reduces yield and makes symptomatic fruit unmarketable.  There are many experimental resources available to the tomato-speck pathosystem, including high-quality tomato and bacterial genomes, CRISPR-Cas9 knockouts of over 150 immunity-associated genes, and many genetic and experimental resources.  We hope our findings in tomato will not only aid tomato improvement, but also serve as a model for other crop species.